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Madagascar: Operation Penguin follows the storyline of the 2005 computer-animated film from the perspective of the intrepid Private Penguin. Players must help Private escape from a hazard-filled zoo and rescue fellow animals from a life of captivity. What ensues is a side-scrolling platform game spanning 18 levels, taking players from monkey cages and lion pens to subways and African plains. Private Penguin can run, jump, swim, and perform his trademark karate chop on such beastly foes as baboons, crocodiles, and snakes. Gadgets can be acquired to assist in the cross-continental journey, including hamburger-wrapper parachutes and soda-bottle jet packs. Movie characters Alex, Marty, Melman, and Gloria are also playable in specific stages tailored to their distinct abilities.